There were actually four times as many bird pairs, or couples, living in the stream-edge habitat compared to the area away from the stream.
为什么只有一个as?
因为这里用了compared to 相当于第二个as. 补全:There were actually four times as many bird pairs, or couples, living in the stream-edge habitat compared to (the bird pairs or couples living in ) the area away from the stream.
 
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