话题 take pleasure in

话题 three runs in

话题 in the hotel

话题 when in walks

话题 in our block

话题 in the table

话题 in some sense

话题 in some senses

话题 in bed awake

话题 in centuries past

话题 in past centuries

话题 in their context

话题 in which direction

话题 in US dollar

问题 关于一个单词then的奇怪用法

...re-introduce wolves into Yellowstone, so that they might usefully eat some of the elk then lumbering about in over-large herds.翻译是这样的：近半个世纪前，生物学家第一次向国会提出重新在黄石引入狼群以便吃掉过度繁殖的麋鹿。主要搞不懂 then 在这个...